This page includes links to PDFs of most of the lab's publications (lab members in bold). Articles are listed in chronological order. Published abstracts and articles in press are not listed. Reprints may be downloaded for personal use by clicking on the icon beside each reference. In doing so, you agree to use reprints exclusively "for purposes such as criticism, comment, news reporting, teaching (including multiple copies for classroom use), scholarship, or research" (United States copyright law, Title 17, United States Code, Section 107).







Yovanovich C.A.M., Grant T., Kelber A. 2019. Differences in ocular media transmittance in classical frog and toad model species and its impact on visual sensitivity. The Journal of Experimental Biology 222: jeb204271.

Yovanovich C.A.M., Pierotti M.E.R., Rodrigues M.T., Grant T. 2019. A dune with a view: the eyes of a neotropical fossorial lizard. Frontiers in Zoology 16: 17.

Rada M., Dias P.H.D.S., Pérez-Gonzalez J.L., Anganoy-Criollo M., Rueda-Solano L.A., Pinto-E M.A., Quintero L.M., Vargas-Salinas F., Grant T. 2019. The poverty of adult morphology: Bioacoustics, genetics, and internal tadpole morphology reveal a new species of glassfrog (Anura: Centrolenidae: Ikakogi) from the Sierra Nevada de Santa Marta, Colombia. PLOS ONE 14: e0215349.

Araujo‐Vieira K., Blotto B.L., Caramaschi U., Haddad C.F.B., Faivovich J., Grant T. 2019. A total evidence analysis of the phylogeny of hatchet-faced treefrogs (Anura: Hylidae: Sphaenorhynchus). Cladistics.

Jeckel A.M., Caorsi V.Z., Grant T., Borges-Martins M. 2019. The nuptial pads of Melanophryniscus (Anura: Bufonidae), with the unexpected occurrence of nuptial-pad–like structures in females of two species. Journal of Herpetology 53: 53–61.

Henrique R.S., Grant T. 2019. Influence of environmental factors on short-term movements of butter frogs (Leptodactylus latrans). Herpetologica 75: 38–46.

Dias P.H.D.S., Brandão A.P., Grant T. 2018. The buccopharyngeal morphology of the tadpole of Ameerega flavopicta (Anura: Dendrobatidae: Colostethinae), with a redescription of its external morphology. Herpetologica 74: 323–328.

Machado D.J., Janies D., Brouwer C., Grant T. 2018. A new strategy to infer circularity applied to four new complete frog mitogenomes. Ecology and Evolution 8: 4011–4018.

Araujo-Vieira K., Caramaschi U., Grillitsch H., Grant T., Faivovich J. 2018. On the identity of Sphaenorhynchus platycephalus (Werner, 1894) (Anura: Hylidae). South American Journal of Herpetology 13:73–84.

Dias P.H.S., Anganoy-Criollo M., Guayasamin J.M., Grant T. 2018. The tadpole of Epipedobates darwinwallacei Cisneros-Heredia and Yánez-Muñoz, 2011 (Dendrobatidae: Colostethinae), with new synapomorphies for Epipedobates. South American Journal of Herpetology 13:54–63.

Faivovich J., Pereyra M.O., Luna M.C., Hertz A., Blotto B.L., Vásquez-Almazán C.R., McCranie J.R., Sánchez D.A., Baêta D., Araujo-Vieira K., Köhler G., Kubicki B., Campbell J.A., Frost D.R., Wheeler W.C., Haddad C.F.B. 2018. On the monophyly and relationships of several genera of Hylini (Anura: Hylidae: Hylinae), with comments on recent taxonomic changes in hylids. South American Journal of Herpetology 13:1–32.

Dias P.H.S., Mongin-Aquino M., Candioti F.V., Carvalho-e-Silva A.M.P.T., Baêta D. 2018. Internal larval morphology of two species of shining leaf frogs (Anura: Phyllomedusidae: Phasmahyla). South American Journal of Herpetology 13:44–53.

Saporito R.A., Grant T. 2018. Comment on Amézquita et al. (2017) “Conspicuousness, color resemblance, and toxicity in geographically diverging mimicry: The pan-Amazonian frog Allobates femoralis.” Evolution 72: 1009–1014. doi:10.1111/evo.13468.

Sánchez-Pacheco S.J., Torres-Carvajal O., Aguirre-Peñafiel V., Sales P.M., Verrastro L., Rivas G.A., Rodrigues M.T., Grant T., Murphy R.W. 2018. Phylogeny of Riama (Squamata: Gymnophthalmidae), impact of phenotypic evidence on molecular datasets, and the origin of the Sierra Nevada de Santa Marta endemic fauna. Cladistics 260–291.

Dias P.H.S. 2018. The tadpole of Proceratophrys bigibbosa (Peters, 1872) (Anura: Odontophrynidae), with a description of its buccopharyngeal morphology and proposal of novel synapomorphies for the P. bigibbosa species group. Copeia 2018: 86–93.

Beneti J.S., Montesinos R., Giovannetti V. (Eds.) 2017. Tópicos de Pesquisa em Zoologia. Instituto de Biociências, São Paulo.

Beneti J.S., Montesinos R., Targino M. 2017. Sistemática filogenética baseada em dados moleculares. Pp. 84–102 in Beneti J.S., Montesinos R., Giovannetti V. (Eds.) 2017. Tópicos de Pesquisa em Zoologia. Instituto de Biociências, São Paulo.

Dias P.H.S., Audino J. 2017. Ontogenia e heterocronia. Pp. 43–53 in Beneti J.S., Montesinos R., Giovannetti V. (Eds.) 2017. Tópicos de Pesquisa em Zoologia. Instituto de Biociências, São Paulo.

Gainett G., Dias P.H.S., Montesinos R. 2017. Metodologia da inferência filogenética. Pp. 67–83 in Beneti J.S., Montesinos R., Giovannetti V. (Eds.) 2017. Tópicos de Pesquisa em Zoologia. Instituto de Biociências, São Paulo.

Henrique R.S., Tiseo G.R., Hernández S.G. 2017. Comportamento animal. Pp. 103–125 in Beneti J.S., Montesinos R., Giovannetti V. (Eds.) 2017. Tópicos de Pesquisa em Zoologia. Instituto de Biociências, São Paulo.

Jeckel A. M., Henrique R.S. 2017. A Zoologia e seu papel na sociedade. Pp. 177–184 in Beneti J.S., Montesinos R., Giovannetti V. (Eds.) 2017. Tópicos de Pesquisa em Zoologia. Instituto de Biociências, São Paulo.

Ospina-Sarria J.J., Cabra-García J. 2017. Parsimony analysis of unaligned sequence data: some clarifications. Cladistics doi:10.1111/cla.12229

Anganoy-Criollo M., Cepeda-Quilindo B. 2017. Redescription of the tadpoles of Epipedobates narinensis and E. boulengeri (Anura: Dendrobatidae). Phyllomedusa 16: 155–182.

Dias I.R., Haddad C.F.B., Argôlo A.J.S., Orrico V.G.D. 2017. The 100th: An appealing new species of Dendropsophus (Amphibia: Anura: Hylidae) from northeastern Brazil. PLOS ONE 12: e0171678.

Enciso-Calle M.P., Viuche-Lozano A., Anganoy-Criollo M., Bernal M.H. 2017. Rediscovery of Atelopus subornatus Werner, 1899 (Anura: Bufonidae), with a redescription of the tadpole. Zootaxa 4344: 160–162.

Mailho-Fontana P.L., Antoniazzi M.M., Rodrigues I., Sciani J.M., Pimenta D.C., Brodie E.D., Rodrigues M.T., Jared C. 2017. Parotoid, radial, and tibial macroglands of the frog Odontophrynus cultripes: Differences and similarities with toads. Toxicon 129: 123–133.

Molina-Zuluaga C., Cano E., Restrepo A., Rada M., Daza J.M. 2017. Out of Amazonia: the unexpected trans-Andean distribution of Cochranella resplendens (Lynch and Duellman, 1978) (Anura: Centrolenidae). Zootaxa 4238: 268–274.

Grant T., Rada M, Anganoy-Criollo M., Batista A., Dias P.H., Jeckel A.M., Machado D.J., Rueda-Almonacid J.V. 2017. Phylogenetic systematics of dart-poison frogs and their relatives revisited. South America Journal of Herpetology 12: S1–S90.

Rada M., Ospina-Sarria J.J., Guayasamin J.M. 2017. A taxonomic review of tan-brown glassfrogs (Anura: Centrolenidae), with the description of a new species from southwestern Colombia. South American Journal of Herpetology 12: 136–156.

Rada M., Jeckel A.M., Caorsi V.Z., Barrientos L.S., Rivera-Correa M., Grant T. 2017. A remarkable new white-eyed glassfrog species of Sachatamia from Colombia (Anura: Centrolenidae), with comments on the systematics of the genus. South American Journal of Herpetology 12: 157–173.

Rivera-Correa M., Vargas-Salinas F., Grant T. 2017. Statistical differences and biological implications: a comparative analysis of the advertisement calls of two Andean stream treefrogs (Hylidae: Hyloscirtus) and the evolution of acoustic characters. Salamandra 53: 237–244.

Blotto B.L., Pereyra M.O., Faivovich J., Dias P.H.S., Grant T. 2017. Concentrated evolutionary novelties in the foot musculature of Odontophrynidae (Anura: Neobatrachia), with comments on adaptations for burrowing. Zootaxa 4258: 425–442.

González-Durán G.A., Targino M., Rada M., Grant T. 2017. Phylogenetic relationships and morphology of the Pristimantis leptolophus species group (Amphibia: Anura: Brachycephaloidea), with the recognition of a new species group in Pristimantis Jiménez de la Espada, 1870. Zootaxa 4243:42–74.

Medeiros C.I., Both C., Grant T., Hartz S.M. 2017. Invasion of the acoustic niche: variable responses by native species to invasive American bullfrog calls. Biological Invasions 19: 675–690.

Galvis P.A., Caorsi V.Z., Sánchez-Pacheco S.J., Rada M. 2016. The advertisement calls of three hylid frogs from Hispaniola. Bioacoustics 25: 89–97.

Segalla M.V., Caramaschi U., Cruz C.A.G., Grant T., Haddad C.F.B., Garcia P.C.A., Berneck B.V.M., Langone J.A. 2016. Brazilian amphibians: list of species. Herpetologia Brasileira 5: 34–46.

Pereyra M.O., Womack M.C., Barrionuevo J.S., Blotto B.L., Baldo D., Targino M., Ospina-Sarria J.J., Guayasamin J.M., Coloma L.A., Hoke K.L., Grant T., Faivovich J. 2016. The complex evolutionary history of the tympanic middle ear in frogs and toads (Anura). Scientific Reports 6: 34130.

Grant T., Frost D.R. 2016. Recent progress in the systematics of poison frogs and their relatives (Dendrobatoidea). Pp. 9–18, in Kahn T. R., La Marca E., Lötters S., Brown J. L., Twomey E., Amézquita A. (Eds.), Aposematic Poison Frogs (Dendrobatidae) of the Andean Countries: Bolivia, Colombia, Ecuador, Perú and Venezuela. Conservation International Tropical Field Guide Series, Conservation lnternational, Arlington.

Rada M., Sánchez-Pacheco J., Brown J.L. 2016. Amazonian poison frog Ranitomeya amazonica (Shulte, 1999). Pp. 441–445, in Kahn T. R., La Marca E., Lötters S., Brown J. L., Twomey E., Amézquita A. (Eds.), Aposematic Poison Frogs (Dendrobatidae) of the Andean Countries: Bolivia, Colombia, Ecuador, Perú and Venezuela. Conservation International Tropical Field Guide Series, Conservation lnternational, Arlington.

Velásquez-Álvarez A.A., Rada M., Kahn T.K. 2016. Dale Swanson ́s poison frog Andinobates daleswansoni (Rueda- Almonacid, Rada, Sánchez-Pacheco, Velásquez-Álvarez and Quevedo, 2006). Pp. 263–267, in Kahn T. R., La Marca E., Lötters S., Brown J. L., Twomey E., Amézquita A. (Eds.), Aposematic Poison Frogs (Dendrobatidae) of the Andean Countries: Bolivia, Colombia, Ecuador, Perú and Venezuela. Conservation International Tropical Field Guide Series, Conservation lnternational, Arlington.

López-López F.J., Rada M., Kahn T.R. 2016. La Planada poison frog Ameerega andina (Myers and Burrowes, 1987). Pp. 116–120, in Kahn T. R., La Marca E., Lötters S., Brown J. L., Twomey E., Amézquita A. (Eds.), Aposematic Poison Frogs (Dendrobatidae) of the Andean Countries: Bolivia, Colombia, Ecuador, Perú and Venezuela. Conservation International Tropical Field Guide Series, Conservation lnternational, Arlington.

Sánchez-Pacheco S.J., Renjifo C., Rada M., Kahn T.R. 2016. Phantasmal poison frog Epipedobates tricolor (Boulenger, 1899). Pp. 362–363, in Kahn T. R., La Marca E., Lötters S., Brown J. L., Twomey E., Amézquita A. (Eds.), Aposematic Poison Frogs (Dendrobatidae) of the Andean Countries: Bolivia, Colombia, Ecuador, Perú and Venezuela. Conservation International Tropical Field Guide Series, Conservation lnternational, Arlington.

Machado D.J., Lyra M.L., Grant T. 2016. Mitogenome assembly from genomic multiplex libraries: Comparison of strategies and novel mitogenomes for five species of frogs. Molecular Ecology Resources 16: 686–693.

Rivera-Correa M., García-Burneo K., Grant T. 2016. A new red-eyed of stream treefrog of Hyloscirtus (Anura: Hylidae) from Peru, with comments on the taxonomy of the genus. Zootaxa 4061: 29–40.

Pereyra M.O., Baldo D., Iglesias P.P., Blotto B.L., Thomé M.T.C., Haddad C.F.B., Barrio-Amorós C.L., Ibáñez R., Faivovich J. 2016. Phylogenetic relationships of toads of the Rhinella granulosa group Anura: Bufonidae): a molecular perspective with comments on hybridization and introgression. Cladistics 32: 36–53.

Barrasso D.A., Alcalde L., Blotto B.L., Basso N.G. 2016. Description of the tadpole of Alsodes neuquensis Cei, 1976 and comparison with the sibling species A. gargola Gallardo, 1970 (Amphibia, Anura, Alsodidae). Herpetological Journal 26:21–31.

Carvalho A.L.G., Sena M.A., Peloso P.L. V, Machado F.A., Montesinos R., Silva H.R., Campbell G., Rodrigues M.T. 2016. A new Tropidurus (Tropiduridae) from the semiarid Brazilian Caatinga: evidence for conflicting signal between mitochondrial and nuclear loci affecting the collared lizards. American Museum Novitates 3852: 1–66.

Araujo-Vieira K., Valdujo P.H., Faivovich J. 2016. A new species of Scinax Wagler (Anura: Hylidae) from Mato Grosso, Brazil. Zootaxa 4061:261–273.

Peloso P.L.V., Orrico V.Z., Haddad C.F.B., Lima-Filho G.R., Sturaro M.J. 2016. A new species of clown tree frog, Dendropsophus leucophyllatus species group, from Amazonia (Anura, Hylidae). South American Journal of Herpetology 11: 66–80.


Ospina-Sarria J.J., Angarita-Sierra T., Pedroza-Banda R. 2015. A new species of Craugastor (Anura: Craugastoridae) from the Magdalena River Valley, Colombia, with evaluation of the characters used to identify species of the Craugastor fitzingeri group. South American Journal of Herpetology 10:143–156.

Jeckel A.M., Saporito R.A., Grant T. 2015. The relationship between poison frog chemical defenses and age, body size, and sex. Frontiers in Zoology 12:27.

Machado D.J. 2015. YBYRÁ facilitates comparison of large phylogenetic trees. BMC Bioinformatics 16:204.

Ospina-Sarria J.J., Velásquez-Trujillo D.A., Bolívar-G. W. 2015. First records of the two poorly known terrarana frogs Pristimantis esmeraldas (Guayasamin, 2004) and Strabomantis necerus (Lynch, 1975) (Amphibia: Anura: Craugastoridae) for Colombia. Herpetology Notes 8:27–30.

Jeckel A.M., Grant T., Saporito R.A. 2015. Sequestered and synthesized chemical defenses in the poison frog Melanophryniscus moreirae. Journal of Chemical Ecology.

Fouquet A., Orrico V.Z., Ernst R., Blanc M., Martinez Q., Vacher J.P., Rodrigues M.T., Ouboter P., Jairam R., Ron S. 2015. A new Dendropsophus Fitzinger, 1843 (Anura: Hylidae) of the parviceps group from the lowlands of the Guiana Shield. Zootaxa 4052:39–64.

Fouquet A., Souza S.M., Nunes P.M.S, Kok P.J.R., Curcio F.F., Carvalho C.M., Grant T., Rodrigues M.T. 2015. Two new endangered species of of Anomaloglossus (Anura: Aromobatidae) from Roraima State, northern Brazil. Zootaxa 3926: 191–210.

Araujo-Vieira K., Brandão R.A., Faria D.C.C. 2015. A new species of rock-dwelling Scinax Wagler (Anura: Hylidae) from Chapada dos Veadeiros, Central Brazil. Zootaxa 3915:52–66.

Araujo-Vieira K., Tacioli A., Faivovich J., Orrico V.G.D., Grant T. 2015. The tadpole of Sphaenorhynchus caramaschii, with comments on larval morphology of Sphaenorhynchus (Anura: Hylidae) Zootaxa 3904: 270–282.

Faivovich J., Nicoli L., Blotto B.L., Pereyra M.O., Baldo D., Barrionuevo J.S., Fabrezi M., Wild E.R., Haddad C.F.B. 2014. Big, bad, and beautiful: Phylogenetic relationships of the horned frogs (Anura: Ceratophryidae). South American Journal of Herpetology 9: 207–227.

Orrico V.Z., Peloso P.L.V., Sturaro M.J., Silva-Filho H.F.D.A., Neckel-Oliveira S., Gordo M., Faivovich J., Haddad C.F.B. 2014. A new “bat-voiced” species of Dendropsophus Fitzinger, 1843 (Anura, Hylidae) from the Amazon Basin, Brazil. Zootaxa 3881: 341–361.

Caorsi V.Z., Colombo P., Freire M.D., Amaral I.B., Zank C., Borges-Martins M., Grant T. 2014. Natural history, coloration pattern and conservation status of the threatened south Brazilian red bellied toad, Melanophryniscus macrogranulosus Braun, 1973. Herpetology Notes 7: 585–598.

de Sá R.O., Grant T., Camargo A., Heyer W.R., Ponssa M.L., Stanley E. 2014. Systematics of the Neotropical genus Leptodactylus( Fitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the relevance of non-molecular evidence, and species accounts. South American Journal of Herpetology 9: S1–S128.

Blotto B.L., Pereyra M.O., Baldo D. 2014. The tadpole of Rhinella azarai (Gallardo, 1965) with comments on larval morphology in the Rhinella granulosa species group (Anura: Bufonidae). Journal of Herpetology 48:434–438.

Gehara M., Crawford A.J., Orrico V.G.D., Rodríguez A., Lötters S., Fouquet A., Barrientos L.S., Brusquetti F., De la Riva I., Ernst R., Urrutia G.G., Glaw F., Guayasamin J.M., Hölting M., Jansen M., Kok P.J.R., Kwet A., Lingnau R., Lyra M., Moravec J., Pombal Jr J.P., Rojas-Runjaic F.J.M., Schulze A., Señaris C.J., Solé M., Rodrigues M.T., Twomey E., Haddad C.F.B., Vences M., Köhler J. 2014. High levels of diversity uncovered in a widespread nominal taxon: continental phylogeography of the Neotropical tree frog Dendropsophus minutus. PLoS One 9:e103958.

Segalla M.V., Caramaschi U., Cruz C.A.G., Grant T., Haddad C.F.B., Langone J.A., Garcia P.C.A. 2014. Brazilian amphibians: list of species. Herpetologia Brasileira 3: 37–48.

Padial J.M., Grant T., Frost D.R. 2014b. Corrections to “Padial et al. (2014) Molecular systematics of terraranas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria”. Zootaxa 3827: 599–600.

Padial J.M., Grant T., Frost D.R. 2014a. Molecular systematics of terraranas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa 3825: 1–132.

Both C., Madalozzo B., Lingnau R., Grant T. 2014. Amphibian richness patterns in Atlantic Forest areas invaded by American bullfrogs. Austral Ecology 39: 864–874.

Zina J., Silva G.R., Loebmann D., Orrico V.G.D. 2014. The recognition of Dendropsophus minusculus (Rivero, 1971) (Hylidae, Dendropsophini) as a highly polymorphic, multi-domain distributed species. Brazilian Journal of Biology 74:146–153.

Grant T., Bolívar-G W. 2014. A new species of semiarboreal toad with a salamander-like ear (Anura: Bufonidae: Rhinella). Herpetologica 70: 198–210.

Dias P.H.S., Carvalho-e-Silva A.M.P.T., Carvalho-e-Silva S.P. 2014. The tadpole of Proceratophrys izecksohni ( Amphibia : Anura : Odontophrynidae ). Zoologia. 31:181–194.

Bolívar-G W., Antoniazzi M.M., Grant T., Jared C. 2014. Discovery of a novel accessory structure of the pitviper infrared receptor organ (Serpentes: Viperidae). PLoS One 9: e90622.

Carvalho M.R., Ebach M.C., Williams D.M., Nihei S.S., Rodrigues M.T., Grant T., Silveira L.F., Gill A.C., Schelly R.C., Sparks J.S., Bockmann F.A., Séret B., Ho H.-C., Grande L., Rieppel O., Dubois A., Ohler A., Faivovich J., Assis L.C.S., Wheeler Q.D., Goldstein P.Z., Almeida E.A.B. de, Valdecassas A.G., Nelson G.J. 2014. Does counting species count as taxonomy? On misrepresenting systematics, yet again. Cladistics 30: 322–329.

Vargas-Salinas F., Quintero-Ángel A., Osorio-Domínguez D., Rojas-Morales J.A., Escobar-Lasso S., Gutiérrez-Cárdenas P.D.A., Rivera-Correa M., Amézquita A., 2014. Breeding and parental behaviour in the glass frog Centrolene savagei (Anura: Centrolenidae). Journal of Natural History 48: 1–17.

Carvajalino-Fernández J.M., Jeckel A.M., Indicatti R.P., 2013. Melanophryniscus moreirae (Amphibia, Anura, Bufonidae): Dormancy and hibernacula use during cold season. Herpetologia Brasileira 2: 61–62.

Grant T., Myers C.W. 2013. Review of the frog genus Silverstoneia, with descriptions of five new species from the Colombian Chocó (Dendrobatidae: Colostethinae). American Museum Novitates 3784: 1–58.

Hantak M.M., Grant T., Reinsch S., McGinnity D., Loring M., Toyooka N., Saporito R.A. 2013. Dietary alkaloid sequestration in a poison frog: an experimental test of alkaloid uptake in Melanophryniscus stelzneri (Bufonidae). Journal of Chemical Ecology 39: 1400–1406.

Taboada C., Grant T., Lynch J.D., Faivovich J. 2013. New morphological synapomorphies for the New World direct-developing frogs (Amphibia: Anura: Terrarana). Herpetologica 69: 342–357.

Díaz-Gutiérrez N., Vargas-Salinas F., Rivera-Correa M., Rojas-Morales J.A., Escobar-Lasso S., Velasco J.A., Gutiérrez-Cárdenas P.D.A., Amézquita A. 2013. Description of the previously unknown advertisement call and tadpole of the Colombian endemic glassfrog Centrolene savagei (Anura: Centrolenidae). Zootaxa 3686: 289–296.

Berneck B., Targino M., Garcia P.C.A. 2013. Rediscovery and re-description of Ischnocnema nigriventris (Lutz, 1925)(Anura: Terrarana: Brachycephalidae). Zootaxa 3694: 131–142.

Cassini C.S., Orrico V.G.D., Dias I.R., Solé M., Haddad C.F.B. 2013. Phenotypic variation of Leptodactylus cupreus Caramaschi, São-Pedro and Feio, 2008 (Anura, Leptodactylidae). Zootaxa 3616: 73–84.

Orrico V.G.D., Duellman W.E., Souza M.B., Haddad C.F.B., 2013. The taxonomic status of Dendropsophus allenorum and Dendropsophus timbeba (Anura: Hylidae). Journal of Herpetology 47: 615–618.

Rivera-Correa M., Duarte-Cubides F., Rueda-Almonacid J.V., Daza J.M. 2013. A new red-eyed treefrog of Agalychnis (Anura: Hylidae: Phyllomedusinae) from middle Magdalena River valley of Colombia with comments on its phylogenetic. Zootaxa 3636: 85–100.

Rivera-Correa M., Orrico V.G.D. 2013. Description and phylogenetic relationships of a new species of treefrog of the Dendropsophus leucophyllatus group (Anura: Hylidae) from the Amazon basin of Colombia and with an exceptional color pattern. Zootaxa 3686: 447–460.

Rivera-Correa M., Faivovich J. 2013. A new species of Hyloscirtus (Anura: Hylidae) from Colombia, with a rediagnosis of Hyloscirtus larinopygion (Duellman, 1973). Herpetologica 69: 298–313.

Suárez P., Cardozo D., Baldo D., Pereyra M.O., Faivovich J., Orrico, V.G.D., Catroli, G.F., Grabiele, M., Bernarde, P.S., Nagamachi, C.Y., Haddad, C.F.B., Pieczarka, J.C. 2013. Chromosome evolution in dendropsophini (Amphibia, Anura, Hylinae). Cytogenetic and Genome Research 141: 295–308.

Caramaschi U., Orrico V.G.D., Faivovich J., Dias I.R., Solé M. 2013. A new species of Allophryne (Anura: Allophrynidae) from the Atlantic rain forest biome of eastern Brazil. Herpetologica 69: 480–491.

Gutiérrez-Cárdenas P.D.A., Rivera-Rojas A., Rivera-Correa M. 2013. Dendropsophus norandinus (north Andean treefrog. Diet. Herpetological Review 44: 120–121.

Both C., Grant T. 2012. Biological invasions and the acoustic niche: the effect of bullfrog calls on the acoustic signals of white-banded tree frogs. Biology Letters 8: 714–716.

Caorsi V.Z., Santos R.R., Grant T. 2012. Clip or Snap? An evaluation of toe-clipping and photo-identification methods for identifying individual southern red-bellied toads, Melanophryniscus cambaraensis. South American Journal of Herpetology 7: 79–84.

Grant T., Colombo P., Verrastro L., Saporito R.A. 2012. The occurrence of defensive alkaloids in non-integumentary tissues of the Brazilian red-belly toad Melanophryniscus simplex (Bufonidae). Chemoecology 22: 169–178.

Grant T. Martins M. 2012. Apresentação da Herpetologia Brasileira. Herpetologia Brasileira 1: 20-21.

Myers C.W., Ibáñez D R., Grant T., Jaramillo C.A. 2012. Discovery of the frog genus Anomaloglossus in Panama, with descriptions of two new species from the Chagres Highlands (Dendrobatoidea: Aromobatidae). American Museum Novitates 3763: 1–20.

Peloso P.L.V., Faivovich J., Grant T., Gasparini J.L., Haddad C.F.B. 2012. An extraordinary new species of Melanophryniscus (Anura, Bufonidae) from southeastern Brazil. American Museum Novitates 3762: 1–31.

Coelho I.P., Teixeira F.Z., Colombo P., Coelho A.V.P., Kindel A. 2012. Anuran road-kills neighboring a peri-urban reserve in the Atlantic Forest, Brazil. Journal of Environmental Management 112: 17–26.

Rivera-Correa M., Gutiérrez-Cárdenas P.D.A. 2012. A new highland species of treefrog of the Dendropsophus columbianus group (Anura: Hylidae) from the Andes of Colombia. Zootaxa 3486: 50–62.

Rivera-Correa M. 2012. Colombian amphibians: cryptic diversity and cryptic taxonomy. Froglog 100, 36–37.

Motta A.P., Castroviejo-Fisher S., Venegas P.J., Orrico V.G.D., Padial J.M., 2012. A new species of the Dendropsophus parviceps group from the western Amazon Basin (Amphibia: Anura: Hylidae). Zootaxa 3249: 18–30.

Fouquet A., Loebmann D., Castroviejo-Fisher S., Padial J.M., Orrico V.G.D., Lyra M.L., Roberto I.J., Kok P.J.R., Haddad C.F.B., Rodrigues M.T. 2012. From Amazonia to the Atlantic forest: molecular phylogeny of Phyzelaphryninae frogs reveals unexpected diversity and a striking biogeographic pattern emphasizing conservation challenges. Molecular Phylogenetics and Evolution 65: 547–61.

Both C., Lingnau R., Santos-Jr A., Madalozzo B., Pedrosa L., T. Grant. 2011. Widespread occurrence of the American bullfrog, Lithobates catesbeianus (Shaw, 1802) (Anura: Ranidae), in Brazil. South American Journal of Herpetology 6: 127–134.

Santos R.R., Grant T. 2011. Diel pattern of migration in a poisonous toad from Brazil and the evolution of chemical defenses in diurnal amphibians. Evolutionary Ecology 25: 249–258.

Bravo-Valencia, L., Rivera-Correa M. 2011. A new species of harlequin frog (Bufonidae: Atelopus) with an unusual behavior. Zootaxa 3045: 57–67.


Grant, T., A.G. Kluge. 2010. REP provides meaningful measurement of support across datasets. Molecular Phylogenetics and Evolution 55: 340–342.

Santos, R.R., S.B. Leonardi, Caorsi, V.Z., Grant, T. 2010. Directional orientation of migration in an aseasonal explosive-breeding toad from Brazil. Journal of Tropical Ecology 26: 415–421.

Zank, C., Di-Bernardo, M., Maneyro, R., Colombo, P., Fusinatto, L.A., Fonte, L.F.M., 2010. Spatial and temporal distribution of Pseudis minuta (Anura, Hylidae, Hylinae) and environmental variables related to its reproductive activity in Reserva Biológica do Lami. Iheringia, Série Zoologica 100: 145–150.

Colombo, P., Zank, C., Bühler, D., Schossler, M., Balestrin, R.L., Santos, R.R., Leonardi, S.B., Borges-Martins, M., Verrastro, L., 2010. Anura, Hylidae, Dendropsophus nahdereri (Lutz and Bokermann, 1963): Distribution extension and new state record. Check List 6: 429–431.

Sánchez-Pacheco, S.J., Rueda-Almonacid, J.V., Rada M. 2010. Notes on the occurrence of Riama simotera (Squamata, Gymnophthalmidae) in Colombia. Herpetological Bulletin 113: 11–13.

Rivera-Correa, M. 2010. Amphibia, Centrolenidae, Centrolene antioquiense (Noble, 1920): New records and geographical distribution in Colombia. Check List. 6: 220–221.

Grant, T., D.A. Fabri, M. Rivera-Correa, & M.A. Rada. 2009. Diversidade reprodutiva e sucesso adaptativo: o caso dos anfíbios. Pages 98–111 in (R)Evolução de Darwin (E. A. Jeckel-Neto & D. G. de Souza, eds.). ediPUCRS, Porto Alegre, Brasil.

Grant, T. & A. G. Kluge. 2009. Parsimony, explanatory power, and dynamic homology testing. Systematics and Biodiversity 7: 357–363.

Myers, C. W. & T. Grant. 2009. Anomaloglossus confusus, a new Ecuadorian frog formerly masquerading as ˝Colostethus˝ chocoensis (Dendrobatoidea: Aromobatidae). American Museum Novitates 3659: 1–12.

Santos, R. R., P. Colombo, S. B. Leonardi, C. Zank, M. Schossler, K. Vieira, T. Grant, M. Borges-Martins, & L. Verrastro. 2009. Amphibia, Anura, Cycloramphidae, Proceratophrys bigibbosa (Peters, 1872) and Proceratophrys brauni Kwet and Faivovich, 2001: Distribution extension and new state record. Check List 5: 922–925.

Frost, D. R., T. Grant, J. Faivovich, A. Haas, C. F. B. Haddad, R. Bain, R. O. de Sá, S. C. Donnellan, C. J. Raxworthy, M. Wilkinson, A. Channing, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. Green and W. C. Wheeler. 2008. Is The Amphibian Tree of Life really fatally flawed? Cladistics 24: 385–395.

Grant, T. & A. G. Kluge. 2008. Clade support measures and their adequacy. Cladistics 24:1051–1064.

Grant, T. & A. G. Kluge. 2008. Credit where credit is due: the Goodman-Bremer support index. Molecular Phylogenetics and Evolution 49: 405–406.

Grant, T., A. R. Acosta-Galvis & J. D. Lynch. 2008. A brief overview of the amphibians of Colombia. Pages 103–104 in Threatened Amphibians of the World (S. N. Stuart, M. Hoffman, J. S. Chanson, R. Berridge, N. Cox, P. Ramani & B. Young, eds.). Lynx Edicions, Barcelona, Spain; IUCN, Gland, Switzerland; and Conservation International, Arlington, USA.

Grant, T. and A. G. Kluge. 2007. Ratio of explanatory power (REP): a new measure of group support. Molecular Phylogenetics and Evolution 44: 483–487.

Grant, T., A. Acosta & M. Rada. 2007. A name for the new species of Allobates (Anura: Dendrobatoidea: Aromobatidae) from the Magdalena Valley of Colombia. Copeia 2007: 844–854.

Grant, T. 2007. A new, toxic species of Colostethus (Anura: Dendrobatiae: Colostethinae) from the Cordillera Central of Colombia. Zootaxa 1555: 39–51.

Savage, J. M., C. W. Myers, D. R. Frost and T. Grant. 2007. Dendrobatidae Cope, 1865 (1850) (Amphibia, Anura): proposed conservation. Bulletin of Zoological Nomenclature 64: 255–260.

Barrientos, L. S., P. C. Pulgarín-R., J. M. Botero-C., and T. Grant. 2006. Fotografías vencejo de cuello castaño en Antioquia. Boletín Sociedad Antioqueña de Ornitología 16: 120–123.

Frost, D. R., T. Grant, and J. R. Mendelson III. 2006. Ollotis Cope, 1875 is the oldest name for the genus currently referred to as Cranopsis Cope, 1875 (Anura: Hyloides: Bufonidae). Copeia 2006: 558.

Frost, D. R., T. Grant, J. Faivovich, R. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, S. C. Donnellan, C. J. Raxworthy, M. Wilkinson, A. Channing, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. Green, and W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.

Kluge, A. G., and T. Grant. 2006. From conviction to anti-superfluity: old and new justifications of parsimony in phylogenetic inference. Cladistics 22: 276–288.

Grant, T., D. R. Frost, R. Ibáñez D., C. W. Myers, and J. M. Savage. 2006. Hyloxalus panamensis Dunn, 1933: proposed emendation of spelling from Hyloxalus panamansis (currently Colostethus panamansis; Amphibia, Anura). Bulletin of Zoological Nomenclature 63: 39–41.

Grant, T., D. R. Frost, J. P. Caldwell, R. Gagliardo, C. F. B. Haddad, P. J. R. Kok, B. D. Means, B. P. Noonan, W. Schargel, and W. C. Wheeler. 2006. Phylogenetic systematics of dart-poison frogs and their relatives (Anura: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History 299: 1–262.

Wheeler, W. C., L. Aagesen, C. P. Arango, J. Faivovich, T. Grant, C. A. D'Haese, D. Janies, W. L. Smith, A. Varón, and G. Giribet. 2006. Dynamic Homology and Phylogenetic Systematics: A Unified Approach Using POY. New York: American Museum of Natural History.


Grant, T. and A. G. Kluge. 2005. Stability, sensitivity, science and heurism. Cladistics 21: 597-604.

Grant, T. 2005. The Phylogeny of Poison Dart Frogs (Amphibia: Anura: Dendrobatidae). PhD Thesis, Columbia University, New York, 530pp.

Grant, T. and A. G. Kluge. 2004. Transformation series as an ideographic character concept. Cladistics 20: 23–31.

Grant, T. 2004. On the identities of Colostethus inguinalis (Cope, 1868) and C. panamensis (Dunn, 1933), with comments on C. latinasus (Cope, 1863) (Anura: Dendrobatidae). American Museum Novitates 3444: 1–24.

Grant, T. and A. G. Kluge. 2003. Data exploration in phylogenetic inference: Scientific, heuristic, or neither. Cladistics 19: 379–418.

Grant, T., J. Faivovich and D. Pol. 2003. The perils of 'point-and-click' systematics. Cladistics 19: 276–285.

Dávalos, L.M., R.R. Sears, G. Raygorodetsky, B.L. Simmons, H. Cross, T. Grant, T. Barnes, L. Putzel, and A.L. Porzecanski. 2003. Regulating access to genetic resources under the Convention on Biological Diversity: an analysis of selected case studies. Biodiversity and Conservation 12: 1511–1524.

Grant, T. and M. C. Ardila-Robayo. 2002. A new species of Colostethus (Anura: Dendrobatidae) from the eastern slopes of the Cordillera Oriental of Colombia. Herpetologica 58: 252–260.

Grant, T. 2002. Testing methods: The evaluation of discovery operations in evolutionary biology. Cladistics 18: 94-111.

Sheil, C. A. and T. Grant. 2001. A new species of colubrid snake (Synophis) from western Colombia. Journal of Herpetology 35: 204–209.

Grant, T. and L. O. Rodríguez. 2001. Two new species of frogs of the genus Colostethus (Dendrobatidae) from Peru and a redescription of C. trilineatus (Boulenger, 1883). American Museum Novitates 3355: 1–24.

Frost, D. R., M. T. Rodrigues, T. Grant, and T. A. Titus. 2001. Phylogenetics of the lizard genus Tropidurus (Squamata: Tropiduridae: Tropidurinae): Direct optimization, descriptive efficiency, and sensitivity analysis of congruence between molecular data and morphology. Molecular Phylogenetics and Evolution 21: 352–371.


Porzecanski , A. L. , Grant T. 2000. Are access regulations achieving the goals of the CBD? IUCN Biology 2: 4–5. [Popular article]

Porzecanski, A. L., R. Sears, T. Grant, L. Putzel, L. Dávalos, T. Barnes, H. Cross, G. Raygorodetsky, B. Simmons, and P. Chasek. 1999. Access to genetic resources: an evaluation of the development and implementation of recent regulation and access agreements. Columbia University School of International and Public Affairs Environmental Policy Studies Working Paper 4: 1-101.

Grant, T. 2000 "1999". Una nueva especie de Rhamphophryne (Anura: Bufonidae) de la Cordillera Central de Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 23: 287–292.

Bolívar-G., W., T. Grant, and L. A. Osorio. 1999. Combat behavior in Centrolene buckleyi and other centrolenid frogs. Alytes 16: 77–83.

Lynch, J. D. and T. Grant. 1998. Dying frogs in western Colombia: catastrophe or trivial observation? Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 23: 149–152

Anonymous. 1998. The Global Taxonomy Initiative: Using Systematic Inventories to Meet Country and Regional Needs. Report of the DIVERSITAS/Systematics Agenda 2000 International Workshop, 17-19 September, 1998, American Museum of Natural History, New York.

Grant, T., W. Bolívar-G., and F. Castro. 1998. The advertisement call of Centrolene geckoideum. Journal of Herpetology 32: 452–455.

Grant, T. and F. Castro-Herrera. 1998. The cloud forest Colostethus (Anura, Dendrobatidae) of a region of the Cordillera Occidental of Colombia. Journal of Herpetology 32: 378–392.

Grant, T. 1998. Una nueva especie de Colostethus del grupo edwardsi de Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 22: 423–428.

Grant, T., E. C. Humphrey, and C. W. Myers. 1997. The median lingual process of frogs: a bizarre character of Old World ranoids discovered in South American dendrobatids. American Museum Novitates 3212: 1–40.